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Mananara Nord National Park

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The Precambrian basement of Mananara, dating back 2,750 million years, belongs to the Antongilian system. Following geological metamorphisms that led to this substrate of rocks composed of amphibolites, micaschists, greenschist and schist, granitic intrusions of about 770 million years ago appeared which, together with Neocene alteration and erosion, are responsible for the orthogonal shape of the region's relief with narrowly linear valleys, rock walls, cascades and waterfalls, falling rocky shores and the hydrographic network following the lines of the landscape and valleys. This explains the difficulty of access to monitor the park, the history of linear fragmentation of the forest and the human occupation along the lines of valleys. The density of vegetation and the ability of forest regeneration worsen the problems as they favour the camouflage of offenders.

Due to the presence of the South Equatorial Current coming from South Asia that runs along the east coast to become Madagascar's southward current, this current brings warm surface waters in all seasons and conditions the water temperature to never drop below 18°-20°C. In order to develop, coral reefs require warm, clear water (above 18°C) on a shallow bottom with 30 to 35% salinity, which is the case for the Mananara Nord Marine Park. At the highest sea (during high tide minus the half maximum ~~hight~~ height of a wave) the water is between 25m and 35m deep. The Malagasy east coast is swept throughout the year by the Alizé (trade wind). The average wind speed is 20m/s.

The Mananara Nord National Park consists of a land and marine park that are both rich in biodiversity as well as natural and original beauty.

The marine park is the first created in Madagascar. The site is commonly called Nosy Antafana. It is a set of three islets located 2.5km off [[Sahasoa]]. The largest is Nosy Be or [[Nosy Antafana]], where a ~~fresh water~~ freshwater source is located. The second is [[Nosy Rangontsy]], named after the man whose remains were the first to be deposited there. On this islet are a mangrove swamp, an expanse of brackish water with fish and in the east huge masses of granite. The last island and the smallest is [[Nosy Hely]]. It is a real marine ecosystem with various representative ecological niches. Coral reefs are among the most beautiful and rich in biodiversity of the east coast of Madagascar. The land park represents the eastern eco-region with littoral forests and forest islands. == Objectives of the site == The Mananara Nord National Park was established on 25th July 1989 by Presidential Decree No. 89/216. It is the first as a Biosphere Reserve in Madagascar. The objective of the site is the conservation of biodiversity in the national park (both marine and terrestrial), it must maintain the quality and quantity of biodiversity in the protected area. The national park is also a water reservoir for the region with thirty sources, the most important are: Ambatomilogno, Fotsialanana, Hoalampano, Ivontaka, Mahavo, Manambato Manandriana, Mandemoko, Menatany, Sahantsidy, Saharamy, Sahasatrana, Sahasoa, Sahofiky, Tsihoronina, Vahibe, Vongohely, ... The Mananara Nord Biosphere Reserve is home to a rich flora and fauna. These species require specific conservation actions as they are subject to pressures. They are key elements in the protected area and are referred to as conservation targets. == Conservation targets == The conservation targets of Mananara Nord National Park are: '''* Dense humid forest of low altitude:''' It comprises almost all of the Mananara Nord Biosphere Reserve. Compared to the very small variation of altitude, the richness in micro-mammal is exceptional. '''* Forest island Nosy Antafana:''' For years the island is exposed to an ongoing attack by a colony of ''Rattus rattus'' (Black rat) invading it. '''* Coral reefs:''' The Mananara reefs are among the most beautiful with a great wealth of biodiversity. '''* The species''' * ''Dypsis antanambensis'' - endemic palm tree, critically endangered (grows only in the park, 49 mature individuals in total)* ''Voanioala gerardii'' - a rare plant worldwide, critically endangered* ''Allocebus trichotis'' - [[Hairy-eared dwarf lemur]], never existed in large numbers. It is found in the vicinity of the Mananara River which is its exclusive habitat. == The site and local population == The Mananara Nord National Park is subject to pressures for the '''* terrestrial part:'''* '''Land clearings''', destroying the forest by burning * '''Human occupations''', as the population increases, inward penetration is inevitable. * '''Illegal logging''' of precious woods and by-products.* '''Poaching''', lemurs, small mammals and birds are most attacked. '''* marine part:'''* Octopus fishing with harpoons* Use of small mesh nets* The abundance of rats on Nosy Antafana == Fauna == === The best fauna most visible to tourists === <table width="100%" cellpadding="2" cellspacing="1" border="0"><tr bgcolor="#dccbb3" valign="top"><td>Scientific name</td><td>English name</td><td>Malagasy name</td><td>Description</td><td>Where to find</td><td>When to find</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Indri indri''</td><td>Indri</td><td>Babakoto</td><td>Completely dark fur</td><td>Verezanantsoro Circuit</td><td>All year round</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Varecia variegata''</td><td>Black-and-white ruffed lemur</td><td>Varikandana</td><td>Easy to observe</td><td>Verezanantsoro Circuit</td><td>All year round</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eulemur albifrons ''</td><td>White-fronted brown lemur</td><td>Varikosibe</td><td>Easy to observe</td><td>Verezanantsoro Circuit, Ivontaka Sud Circuit, Varary Circuit</td><td>During the day</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Brookesia superciliaris''</td><td>Brown leaf chameleon</td><td>Ranovary</td><td>Tiny chameleon</td><td>Verezanantsoro Circuit, Ivontaka Sud Circuit, Varary Circuit</td><td>October to November</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Euryceros prevostii''</td><td>[[Helmet vanga]]</td><td>Siketribe</td><td>Indicator of an undisturbed forest</td><td>Verezanantsoro Circuit</td><td>All year round</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Pteropus rufus''</td><td>Madagascan flying fox</td><td>Fanihy</td><td></td><td>Nosy Antafana Circuit</td><td>All year round</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Amphiprion latifasciatus''</td><td>Madagascar anemonefish</td><td></td><td>Indicator of an intact habitat</td><td>Nosy Antafana aquatic Circuit</td><td>All year round</td></tr></table> Inventories have identified: On the terrestrial fauna,* 77 species of birds* 136 species of herpetofauna (reptiles and amphibians of a particular region) identified throughout a survey in 2001* 13 species of lemurs in the forest of the reserve* 17 rodent species known in the low to medium altitude moist forest (0 - 1,200m) in northeastern Madagascar* 7 species of freshwater fish in the surrounding of the park* Regarding arthropods, Mananara Nord National Park forms with [[Zahamena National Park]] and [[Masoala National Park]] what is called the endemic centre of Madagascar* The Mananara Nord National Park is one of the protected areas that has the highest number of micro-mammal species in the eastern ecoregion given its reduced altitude range and size, which barely covers one-tenth of the Masoala surface. Indeed, there are 16 species against 17, for which Masoala is the richest. On the marine fauna,* 132 species of cnidaria* 64 species of molluscs* 16 species of arthropods* 34 species of echinoderms* 179 species of fish* 2 species of reptiles* 2 species of mammals === Lemurs === An inventory identified 13 species of lemurs.<table width="100%" cellpadding="2" cellspacing="1" border="0"><tr bgcolor="#dccbb3" valign="top"><td>Scientific name</td><td>English name</td><td>Malagasy name</td><td>Endemicity</td><td>Description</td><td>Biology and behavior</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Indri indri''</td><td>[[Indri]]</td><td>Babakoto</td><td>Endemic to Madagascar</td><td>Black fur, the largest of the lemurs</td><td>Arboreal (lives on trees), folivore (feeds on leafs), group of 2 to 3</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Propithecus diadema''</td><td>[[Diademed sifaka]]</td><td>Sifaka</td><td>Endemic to Madagascar</td><td>White fur, red thigh, fairly large</td><td>Diurnal, lives in a group</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Avahi laniger''</td><td>[[Eastern woolly lemur]]</td><td>Ampongy</td><td>Endemic to Madagascar</td><td>Grey fur, average size</td><td>Twilight active, lives in a group</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Daubentonia madagascariensis''</td><td>[[Aye-aye]]</td><td>Haihay</td><td>Endemic to Madagascar</td><td>Black fur, average size</td><td>Nocturnal, insectivore (feeds on insects), solitary</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Varecia variegata''</td><td>[[Black-and-white ruffed lemur]]</td><td>Varikandana</td><td>Endemic to Madagascar</td><td>Black and white fur, fairly large</td><td>Diurnal, lives in a group of more than 4</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eulemur albifrons''</td><td>[[White-fronted brown lemur]]</td><td>Varikosibe</td><td>Endemic to Madagascar</td><td>Grey fur, average size</td><td>Diurnal, lives in group of more than 6</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eulemur fulvus''</td><td>[[Brown lemur]]</td><td>Varikosy</td><td>Endemic to Madagascar</td><td>Grey coat, black muzzle, average size</td><td>Diurnal or twilight active, lives in group of up to 12</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eulemur rubriventer''</td><td>[[Red-bellied lemur]]</td><td>Alomena</td><td>Endemic to Madagascar</td><td>Red fur, average size</td><td>Diurnal, lives in group of up to 4</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Hapalemur griseus''</td><td>[[Eastern grey bamboo lemur]]</td><td>Bokombolo</td><td>Endemic to Madagascar</td><td>Grey fur, relatively small size</td><td>Diurnal or twilight active, folivore (feeds on leafs), lives in group of several individuals</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lepilemur mustelinus''</td><td>[[Weasel sportive lemur]]</td><td>Fitsidiky</td><td>Endemic to Madagascar</td><td>Grey fur, medium size</td><td>Nocturnal</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Cheirogaleus major''</td><td>[[Greater dwarf lemur]]</td><td>Tsitsihy</td><td>Endemic to Madagascar</td><td>Grey fur, relatively small size</td><td>Nocturnal, hibernates during winter</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Allocebus trichotis''</td><td>[[Hairy-eared dwarf lemur]]</td><td>Tsidiala</td><td>Endemic to Madagascar</td><td>Dark grey fur, small size</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Microcebus rufus''</td><td>[[Brown mouse lemur]]</td><td>Tsidy</td><td>Endemic to Madagascar</td><td>Grey coat, smallest lemur</td><td>Nocturnal, more insectivore (feeds on insects) than frugivore (feeds on fruits)</td></tr></table> === Birds === There are 77 species of birds in the park. <table width="100%" cellpadding="2" cellspacing="1" border="0"><tr bgcolor="#dccbb3" valign="top"><td>Scientific name</td><td>English name</td><td>Malagasy name</td><td>Endemicity</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Accipiter francesiae''</td><td>Frances's sparrowhawk</td><td>Firasa / Fihiaky</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Accipiter madagascariensis''</td><td>[[Madagascar sparrowhawk]]</td><td>Firasa</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Acridotheres tristis''</td><td>Common myna</td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Agapornis canus''</td><td>[[Grey-headed lovebird]]</td><td>Karaoko</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Alectroenas madagascariensis''</td><td>Madagascar blue pigeon</td><td>Finengo maitso</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Asio madagascariensis''</td><td>Madagascar long-eared owl</td><td>Hankagna</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Brachypteracias leptosomus''</td><td>Short-legged ground-roller</td><td>Fangadiovy</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Brachypteracias squamiger''<br>now ''Geobiastes squamiger''</td><td>Scaly ground-roller</td><td></td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Buteo brachypterus''</td><td>Madagascar buzzard</td><td>Hindry / Beririnina</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Calicalicus madagascariensis''</td><td>Red-tailed vanga</td><td>Vanga mena / Totokarasoroka</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Canirallus kioloides'' or ''Mentocrex kioloides''</td><td>Madagascar wood-rail</td><td>Tasikoza vohitra</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Caprimulgus madagascariensis''</td><td>Madagascar nightjar</td><td>Tataro</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Centropus toulou''</td><td>Madagascar coucal</td><td>Toloho</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Cisticola cherina''</td><td>Madagascar cisticola</td><td>Tity / Tsintsina</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Copsychus albospecularis''</td><td>[[Madagascar magpie-robin]]</td><td>Todiagna</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Coracina cinerea''<br>now ''Ceblepyris cinereus''</td><td>Madagascar cuckoo-shrike</td><td>Vorondavenoky / Vorondavenona</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Coracopsis nigra''</td><td>Black parrot</td><td>Boeza fihitra</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Coracopsis vasa''</td><td>Greater vasa parrot</td><td>Boeza be</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Corythornis vintsioides''</td><td>[[Madagascar kingfisher]]</td><td>Vintsy</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Coua caerulea''</td><td>Blue coua</td><td>Mariha</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Coua cristata''</td><td>Crested coua</td><td>Ambosanga</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Coua reynaudii''</td><td>Red-fronted coua</td><td>Koa</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Coua serriana''</td><td>Red-breasted coua</td><td>Koa</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Cuculus rochii''</td><td>Madagascar cuckoo</td><td>Taotaonkafa</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Cyanolanus madagascarinus''</td><td></td><td></td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Cypsiurus parvus''</td><td>African palm swift</td><td>Firihitsandro / Tsiditsidina</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Dryolimnas cuvieri''</td><td>Dryolimnas cuvieri</td><td>Tsikoza</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Dicrurus forficatus''</td><td>[[Crested drongo]]</td><td>Ledronga</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Euryceros prevostii''</td><td>[[Helmet vanga]]</td><td>Siketribe</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eurystomus glaucurus''</td><td>Broad-billed roller</td><td>Tsararahaky</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Falco newtoni''</td><td>Madagascar kestrel</td><td>Hitsikitsika</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Foudia madagascariensis''</td><td>[[Madagascar red fody]]</td><td>Fody / Fodimena</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Foudia omissa''</td><td>Forest fody</td><td>Fody / Fodiala</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Hartlaubius auratus''</td><td>Hartlaubius auratus</td><td>Vorontainomby</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Hypositta corallirostris''</td><td>Coral-billed nuthatch vanga</td><td>Sakodidy</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Hypsipetes madagascariensis''</td><td>Madagascar bulbul</td><td>Menasony</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Ispidina madagascariensis''<br>now ''Corythornis madagascariensis''</td><td>Madagascar pygmy-kingfisher</td><td>Vintsiala</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Leptopterus chabert''</td><td>Chabert's vanga</td><td>Fotsy tretraka</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Leptopterus viridis''<br>now ''Artamella viridis''</td><td>White-headed vanga</td><td>Vanga</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Leptosomus discolor''</td><td>Cuckoo-roller</td><td>Kirombo</td><td></td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lonchura nana''</td><td>Madagascar mannikin</td><td>Tsingirity</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lophotibis cristata''</td><td>Madagascar crested ibis</td><td>Lampirana</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Margaroperdix madagarensis''</td><td>[[Madagascar partridge]]</td><td>Traotrao</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Merops superciliosus''</td><td>Olive bee-eater</td><td>Kirokirioky</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mesitornis unicolor''</td><td>Brown mesite</td><td>Roa telo, voronantambo</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Motacilla flaviventris''</td><td>[[Madagascar wagtail]]</td><td>Triotrio</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mystacornis crossleyi''</td><td>Crossley’s babbler</td><td></td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Nectarinia notata''</td><td>Long-billed green sunbird</td><td>Soy</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Nectarinia souimanga''or ''Cinnyris sovimanga''</td><td>Souimanga sunbird</td><td>Soy</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Neodrepanis coruscans''</td><td>Sunbird asity</td><td>Soy ala</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Neomixis striatigula''</td><td>Stripe-throated jery</td><td>Jijy</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Neomixis tenella''</td><td>Common jery</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Neomixis viridis''</td><td>Green jery</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Nesillas typica''</td><td>Madagascar brush-warbler</td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Newtonia amphichroa''</td><td>Dark newtonia</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Newtonia brunneicauda''</td><td>Common newtonia</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Newtonia fanovanae''</td><td>Red-tailed newtonia</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Ninox superciliaris''<br>now ''Athene superciliaris''</td><td>[[White-browed hawk-owl]]</td><td>Vorondolo</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Numida meleagris''</td><td>Helmeted guineafowl</td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Oriolia bernieri''</td><td>[[Bernier's vanga]]</td><td>Taporo</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Otus rutilus''</td><td>Madagascar scops-owl</td><td>Torotoroka</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Oxylabes madagascariensis''</td><td>White-throated oxylabes</td><td>Sirontsirona</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Philepitta castanea''</td><td>[[Velvet asity]]</td><td>Asity</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Phyllastrephus madagascariensis''<br>or ''Bernieria madagascariensis''</td><td>Long-billed bernieria</td><td>Tretreky</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Phyllastrephus zosterops''<br>or ''Xanthomixis zosterops''</td><td>Short-billed greenbul</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Ploceus nelicourvi''</td><td>Nelicourvi weaver</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Polyboroides radiatus''</td><td>Madagascar harrier-hawk</td><td>Fihiaka</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Randia pseudozosterops''</td><td>Rand's warbler</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Sarothrura insularis''</td><td>Madagascar flufftail</td><td>Mpangalatrovy</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Schetba rufa''</td><td>[[Rufous vanga]]</td><td>Voronantambo</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Streptopelia picturata''<br>or ''Nesoenas picturatus''</td><td>Madagascar turtle-dove</td><td>Domohina</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Terpsiphone mutata''</td><td>[[Madagascar paradise-flycatcher]]</td><td>Soketry / Tsingitry</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Treron australis''</td><td>Madagascar green pigeon</td><td>Finengo mavo</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Tylas eduardi''</td><td>Tylas vanga</td><td>Kikimavo</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Vanga curvirostris''</td><td>Hook-billed vanga</td><td>Vanga soratra</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Zoonavena grandidieri''</td><td>Madagascar spinetail</td><td>Manaviandro</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Zosterops maderaspatanus''</td><td>[[Malagasy white-eye]]</td><td>Tsiparimaso</td><td>   </td></tr></table> === Tenrecs === There are 9 species of tenrecs in the park. <table width="100%" cellpadding="2" cellspacing="1" border="0"><tr bgcolor="#dccbb3" valign="top"><td>Scientific name</td><td>English name</td><td>Malagasy name</td><td>Endemicity</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Hemicentetes semispinosus''</td><td>Lowland streaked tenrec</td><td>Antsora</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Microgale brevicaudata''</td><td>Short-tailed shrew tenrec</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Microgale dobsoni''</td><td>Dobson's shrew tenrec</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Microgale parvula''</td><td>Pygmy shrew tenrec</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Microgale principula''</td><td>Greater long-tailed shrew tenrec</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Microgale talazaci''</td><td>Talzac's shrew tenrec</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Oryzorictes hova''</td><td>Mole-like rice tenrec</td><td>Antsangy</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Setifer setosus''</td><td>Greater hedgehog tenrec</td><td>Sokina</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Tenrec ecaudatus''</td><td>Tailess tenrec</td><td>Trandraka</td><td>Endemic to Madagascar</td></tr></table> === Rodents === There are 7 rodent species in the park. <table width="100%" cellpadding="2" cellspacing="1" border="0"><tr bgcolor="#dccbb3" valign="top"><td>Scientific name</td><td>English name</td><td>Malagasy name</td><td>Endemicity</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Suncus murinus''</td><td>House shrew</td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eliurus grandidieri''</td><td>Grandidier's tuft-tailed rat</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eliurus minor''</td><td>Lesser tufted-tailed rat</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eliurus myoxinus''</td><td>Western tuft-tailed rat</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eliurus tanala''</td><td>Tanala tuft-tailed rat</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eliurus webbi''</td><td>Webb's tuft-tailed rat</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Nesomys rufus''</td><td>Eastern red forest rat</td><td>   </td><td>Endemic to Madagascar</td></tr></table> === Herpetofauna - reptiles and amphibians === An overview of herpetofauna in the park: <table width="100%" cellpadding="2" cellspacing="1" border="0"><tr bgcolor="#dccbb3" valign="top"><td>Scientific name</td><td>English name</td><td>Malagasy name</td><td>Endemicity</td></tr> <tr bgcolor="#eadece" valign="top"><td colspan="4">'''Frogs'''</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Heterixalus betsileo''</td><td>Betsileo reed frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Heterixalus madagascariensis''</td><td>Madagascar reed frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Heterixalus punctatus''</td><td>   </td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Heterixalus tricolor''</td><td>Three-color reed frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantella betsileo''</td><td>Betsileo golden frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantella nigricans''</td><td>Guibé's mantella</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantella pulchra''</td><td>Parker's golden frog</td><td>   </td><td>Locally endemic</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus aglavei''<br>now ''Spinomantis aglavei''</td><td>Anamalozoatra Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus albofrenatus''</td><td>Eastern Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus ambreensis''</td><td>   </td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus argenteus''</td><td>Folohy Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus asper''<br>now ''Gephyromantis asper''</td><td>East Betsileo Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus betsileanus''</td><td>Betsileo Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus bicalcaratus''<br>now ''Guibemantis bicalcaratus''</td><td>Sainte Marie Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus biporus''</td><td>Two-pore Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus glomersae''</td><td>   </td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus boulengeri''<br>now ''Gephyromantis boulengeri''</td><td>   </td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus cormicus''</td><td>   </td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus femoralis''</td><td>Fort Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus fimbriatus''<br>now ''Spinomantis fimbriatus''</td><td>   </td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus grandidieri''</td><td>Grandidier's Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus grandisonae''<br>now ''Blommersia grandisonae''</td><td>Ambrana Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus granulatus''<br>now ''Gephyromantis granulatus''</td><td>Grainy Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus bicalcaratus''<br>now ''Guibemantis bicalcaratus''</td><td>Sainte Marie Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus clemerii</td><td>   </td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus leucomaculatus''<br>now ''Gephyromantis leucomaculatus ''</td><td>White-spotted Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus liber<br>now ''Guibemantis liber''</td><td>Free Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus lugubris''<br><td>Dumeril's Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus luteus''<br>now ''Gephyromantis luteus''</td><td>White Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus malagasius''<br>now ''Gephyromantis malagasius''</td><td>Malasay Grainy Frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus ocarbii''</td><td>   </td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus opiparis''</td><td>Central Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus terakae''</td><td>   </td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus plucifer''</td><td>Common Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus pseudoasper''<br>now ''Gephyromantis pseudoasper''</td><td>Massif Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus pulcher''<br>now ''Guibemantis pulcher''</td><td>Tsarafidy Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus redimitus''<br>now ''Gephyromantis redimitus''</td><td>   </td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus rivicola''<br>now ''Gephyromantis rivicola''</td><td>   </td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus tornieri''<br>now ''Guibemantis tornieri''</td><td>   </td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Mantidactylus wittei''<br>now ''Blommersia wittei''</td><td>Witte's Madagascar frog</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td colspan="4">'''Geckos'''</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Blaesodactylus antongilensis''</td><td>   </td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Edoneva inensus''</td><td>   </td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Geckolepis maculata''</td><td>Fish-scale gecko</td><td>Fingoko</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Geckolepis typica''</td><td>Grandidier's Gecko</td><td>Fingoko</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Hemidactylus frenatus''</td><td>Common house gecko</td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Hemidactylus mercatorius''</td><td>   </td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lygodactylus guibei''</td><td>Western dwarf gecko</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lygodactylus miops''</td><td>Günther's dwarf gecko</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Paroedura gracilis''</td><td>Graceful Madagascar ground gecko</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Phelsuma dorsivittata''</td><td>   </td><td>Antsasatra</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Phelsuma lineata bifasciata''<br>or ''Phelsuma lineata lineata''</td><td>Lined day gecko</td><td>Antsasatra</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Phelsuma madagascariensis madagascariensis''</td><td>Madagascar day gecko</td><td>Antsasatra</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Phelsuma pusilla pusilla''</td><td>   </td><td>Antsasatra</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Phelsuma serraticauda''</td><td>Flat-tailed day gecko</td><td>Antsasatra</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Uroplatus fimbriatus''</td><td>Common flat-tail gecko</td><td>Antaifisaka</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Uroplatus lineatus''</td><td>Lined flat-tail gecko</td><td>Antaifisaka</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Uroplatus sikorae''</td><td>Mossy leaf-tailed gecko</td><td>Antaifisaka</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td colspan="4">'''Chameleons'''</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Brookesia griveaudi''</td><td>Marojejy leaf chameleon</td><td>Ranovary</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Brookesia peyrierasi''</td><td>Peyrieras's pygmy chameleon</td><td>Ranovary</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Brookesia superciliaris''</td><td>Brown leaf chameleon</td><td>Ranovary</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Calumma gastrotaenia''</td><td>Short-nosed chameleon</td><td>Tandrondro</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Calumma nasutum''</td><td>Nose-horned chameleon</td><td>Tandrondro</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Furcifer lateralis''</td><td>Carpet chameleon</td><td>Taintotro</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Furcifer oustaleti''</td><td>Oustalet's chameleon</td><td>Taintotro</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Furcifer pardalis''</td><td>Panther chameleon</td><td>Taintotro</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td colspan="4">'''Skinks'''</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Amphiglossus anosyensis''</td><td>   </td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Amphiglossus astrolabi''</td><td>Diving Skink</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Amphiglossus frontoparietalis''</td><td>Boulenger's Tree Skink</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Amphiglossus melanopleura''<br>now ''Madascincus melanopleura''</td><td>Common Madagascar Skink</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Amphiglossus ornaticeps''</td><td>Gray Skink</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Trachylepis gravenhorstii''</td><td>Gravenhorst's mabuya</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td colspan="4">'''Lizards'''</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Zonosaurus gregoir''</td><td>   </td><td>Androngo</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Zonosaurus madagascariensis''</td><td>Madagascar girdled lizard</td><td>Androngo</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td colspan="4">'''Snakes'''</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Sanzinia madagascariensis''</td><td>Madagascar tree boa</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Dromicodryas bernieri''</td><td>Bernier's striped snake</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Geodipsas infralineata''<br>now ''Compsophis infralineatus''</td><td>Lined forest snake</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Ithycyphus perineti''</td><td>Perinet's night-snake</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Leioheterodon madagascariensis''</td><td>Malagasy giant hognose snake</td><td>Mantingoro</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Liophidium rhodogaster''</td><td>Gold-collared snake</td><td>Lapata</td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Liopholidophis lateralis''<br>now ''Thamnosophis lateralis''</td><td>Lateral water snake</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Liopholidophis thieli''<br>now ''Thamnosophis infrasignatus''</td><td>Forest water snake</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Madagascarophis colubrinus''</td><td>Madagascar night snake</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Macropisthodon ochraceus''</td><td>   </td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Pseudoxyrhopus heterurus''</td><td>Night brook snake</td><td>Mantsiviry </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Pseudoxyrhopus microps''</td><td>Brown brook snake</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Pelamis platurus''<br>now ''Hydrophis platurus''</td><td>Yellow-bellied sea snake</td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Stenophis arctifasciatus''<br>now ''Phisalixella arctifasciata''</td><td>   </td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Typhlops decorsei''</td><td>Mocquard's worm snake</td><td>   </td><td>Endemic to Madagascar</td></tr> <tr bgcolor="#eadece" valign="top"><td colspan="4">'''Turtles'''</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Caretta caretta''</td><td>Loggerhead turtle</td><td>Fano</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Eretmochelys imbricata''</td><td>Hawksbill turtle</td><td>Fanosisiky</td><td>   </td></tr></table> === Marine life - fish === An overview of the marine life in the park: <table width="100%" cellpadding="2" cellspacing="1" border="0"><tr bgcolor="#dccbb3" valign="top"><td>Scientific name</td><td>English name</td><td>Malagasy name</td><td>Endemicity</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Ablennes hians''</td><td>Flat needlefish</td><td>Antseraka</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Acanthurus xanthopterus''</td><td>Yellowfin surgeonfish</td><td>Henalaza</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Acanthurus triostegus''</td><td>Convict surgeonfish</td><td>Rarinkigny</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Albula glossodonta''</td><td>Shortjaw bonefish</td><td>Fiamandry</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Alepes djedaba''</td><td>Shrimp scad</td><td>Boniky</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Anampses caeruleopunctatus''</td><td>Bluespotted wrasse</td><td>Kalama</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Caesio sp.''</td><td>   </td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Carangoides armatus''</td><td>Longfin trevally</td><td>Lagnaorana</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Cephalopholis sonnerati''</td><td>Tomato hind</td><td>(Bête rouge)</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Chaetodon fasciatus''</td><td>Diagonal butterflyfish</td><td>Tatangy</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Chaetodon lunula''</td><td>Halfmoon butterflyfish</td><td>Tatangy</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Chaetodon trifasciatus''</td><td>Lineated butterflyfish</td><td>Tatangy</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Cheilinus fasciatus''</td><td>Redbreasted wrasse</td><td>Bekoravina</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Cheilinus trilobatus''</td><td>Tripletail wrasse</td><td>Bekoravina</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Cheilio inermis''</td><td>Cigar wrasse</td><td>Laoko ahitra</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Chirocentrus nudus''</td><td>Whitefin wolf-herring</td><td>Laotsabatra</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Coris aygula''</td><td>Clown coris</td><td>Ramihamigny</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Ctenochaetus striatus''</td><td>Striped bristletooth</td><td>Henalaza</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Dascyllus aruanus''</td><td>Whitetail dascyllus</td><td>Tsigoagoaka</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Drepane punctata''</td><td>Spotted sicklefish</td><td>Takobatra</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Echeneis naucrates''</td><td>Whitefin sharksucker</td><td>Amby</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Epinephelus posteli''</td><td>Striped fin rockcod</td><td>Alovo</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Epinephelus tauvina''</td><td>Greasy grouper</td><td>Alovo</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Gerres acinaces''</td><td>Smallscale pursemouth</td><td>Fiampotsy</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Gymnothorax permistus''<br>now ''Gymnothorax favagineus''</td><td>Laced moray</td><td>   </td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Hemiramphus far''</td><td>Black-barred halfbeak</td><td>Antserabandana</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Kyphosus cinerascens''</td><td>Blue sea chub</td><td>Torovoko</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Leiognathus equulus''</td><td>Common ponyfish</td><td>Salelo</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Leptoscarus vaigiensis''</td><td>Marbled parrotfish</td><td>Laoko ahitra</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lethrinus elongatus''<br> now ''Lethrinus microdon''</td><td>Smalltooth emperor</td><td>Ambitry lavaorona</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lethrinus harak''</td><td>Thumbprint emperor</td><td>Mandrantongona</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lethrinus lentjan''</td><td>Pinkear emperor</td><td>Menaheliky</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lethrinus mahsena''</td><td>Sky emperor</td><td>Aleta</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lethrinus nebulosus''</td><td>Spangled emperor</td><td>Ambitry</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lethrinus ramak''<br>now ''Lethrinus obsoletus''</td><td>Orange-striped emperor</td><td>Ambitry fotsy</td><td>   </td></tr> <tr bgcolor="#eadece" valign="top"><td>''Lethrinus rubrioperculatus''</td><td>Spotcheek emperor</td><td>Menavalogno</td><td>   </td></tr></table> == Flora == The best flora most visible to tourists: <table width="100%" cellpadding="2" cellspacing="1" border="0"><tr bgcolor="#dccbb3" valign="top"><td>Scientific name</td><td>English name</td><td>Malagasy name</td><td>Endemicity</td><td>Where to find</td><td>When to find</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Galeola humblotii''</td><td>   </td><td>Lavanioala</td><td>Locally endemic</td><td>Ivontaka Sud Circuit</td><td>All year round</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Dypsis antanambensis''</td><td></td><td>Vontro rano</td><td>Locally endemic</td><td>PK 8 Circuit</td><td>All year round</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Dypsis ramentacea''</td><td>   </td><td>Sinkara</td><td>Locally endemic</td><td>PK 8 Circuit</td><td>All year round</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Satranala decussilvae''</td><td>   </td><td>Satranala</td><td>Locally endemic</td><td>PK 8 Circuit, Ivontaka Sud Circuit, Varary Circuit</td><td>All year round</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Dypsis beentjei''</td><td>   </td><td>   </td><td>Locally endemic</td><td>PK 8 Circuit</td><td>All year round</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Dypsis fanjana''</td><td>   </td><td>Fanjana</td><td>Locally endemic</td><td>PK 8 Circuit</td><td>All year round</td></tr> <tr bgcolor="#eadece" valign="top"><td>''Voanioala sp.''</td><td>   </td><td>Voanioala</td><td>Locally endemic</td><td>Vohitrampongy Circuit, Verezanantsoro Circuit</td><td>All year round</td></tr></table> The Mananara Nord National Park counts: '''On the terrestrial flora,''' * more than 764 primary forest species* 436 modified forest species, totalling more than 1200 species of vascular plants including * 54 Cryptogams (Pteridophytes) - disperse spores* 108 Angiosperms (Monocotyledons) - flowering plants* 861 Angiosperms (Dicotyledons)* 1 Gymnosperm - seed-producing plants. These terrestrial ecosystem species are distributed in approximately * 418 genera* 117 families The families most diverse in species are * 66 species of Rubiaceae - family of flowering plants, commonly known as the coffee family* 50 species of Euphorbiaceae - family of flowering plants, the spurge family* 47 species of Arecaceae - family of perennial plants, commonly known as palms* Fabaceae - family of flowering plants, commonly known as the legume, pea or bean family* Flacourtiaceae - defunct family of flowering plants* 27 species of Sapotaceae - family of flowering plants, evergreen trees and shrubs. The palm group is the particularity of Mananara Nord National Park. It can be seen from sea level to the peak of Verezanantsoro. Currently, there are 47 including at least 2 locally endemic palm species. These are ''Dypsis antanambensis'' and ''Dypsis linearis'' (''Dypsis anovensis''). '''On the marine flora,''' * 114 species in 3 different habitats. The seagrass counts: * 32 species of Algae* 9 species of Phanerogams - seed plants. The mangroves are home to: * 7 out of 9 mangrove species found in Madagascar. On the islands of Nosy Atafana, Nosy Rangontsy and Nosy Hely grow 98 species of plants. Of the 3 types of littoral forest (near sea level on laterite, sand and granite blocks) in the reserve, those of the islands of the marine park are the poorest in species. === Habitats === The '''terrestrial park''' includes a terrestrial ecosystem with two types of plant formations: * low-altitude rainforest* savoka - secondary vegetation made up of different types** ''Psiadia altissima''** ''Trema orientalis''** ''Harungana madagascariensis''** ''Ravenala madagascariensis''** ''Anthostema madagascariensis''** ''Uapaca ferruginea''. These different types of formation grow on the ground generally ferralitic and rest on the crystalline base: granite of Antongil and migmatite of Mananara. In addition to these two types of formations encountered in the park, there are also the coastal forests growing along the coast and mainly between Antanambe and Anove. In the '''marine park''' following types of environment are distinguished: * mangroves* grass beds* inner flats* external slopes,* rocky cliffs* passes. Among these environments are the inner flats and the external slopes north, south and west which are the richest in fish species.

== How to get there ==

'''By boat''' from Mananara to the park (1-1.5 hour by speedboat, 3-4 hours by slow boat), or from [[Antanambe]] to the park (45 minutes by speedboat). From Soanierana Ivongo or [[Sainte Marie]] to Mananara it takes 5 hours by boat respectively. The sea is calm from April to October.

== Guides and entrance fees ==

There are two park offices located in Antanambe and Mananara (Mahambolona). There you can get all the information regarding guides, tours, admission fees within the park and guide rates.

There are several guides available for your visit to the park. They all speak French and English and have received training in guiding techniques and flora and fauna.

The admission fee for visiting the park for foreign tourists is 45,000 Ariary per day, children pay 25,000 Ariary per day (prices 2015). Guides are obligatory. The fee for a guide depends on the chosen tour (circuit). There are also porters available. The fee for a porter is 12,000 Ariary per day, including his meals and accommodation.

== Circuits (Tours) ==

=== Verezanantsoro Circuit ===

Medium circuit, 20km in one day. Sudden climate change: rain, temperature difference between sunny area and in dense forest area.

'''What to see?'''<br>

You will cross cloves and vanilla plantations in a beautiful landscape along the coast, amazing panoramic view before diving into the heart of a low-altitude forest with exceptional biodiversity such as lemurs (Indri, Black-and-white ruffed lemur), birds…<br>